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However, the methods from the aforementioned applications cannot be directly applied to the nephron tracking problem due to a number of factors. A crucial difference is that there are hundreds to thousands of nephrons [15] that need to be independently tracked through serial slices (a three-dimensional problem) as opposed to one or a few structures in single images (a two-dimensional problem). In particular, the tortuosity of the nephrons poses a major challenge. Nevertheless, several concepts from existing tracking applications have been adopted in the current approach, such as graph-based tracking, metrics to indicate confidence per iteration, and a set of validation rules to reduce error. 2b1af7f3a8